Ake T. Lu1Department of Human Genetics, David Geffen School of Medicine, University of California Los Angeles, Los Angeles, CA 90095, USA, Anne Seeboth2Centre for Genomic and Experimental Medicine, Institute of Genetics & Molecular Medicine, University of Edinburgh, Edinburgh EH4 2XU, UK, Pei-Chien Tsai3,Department of Twin Research and Genetic Epidemiology, Kings College London, London SE1 7EH, UK4,Department of Biomedical Sciences, Chang Gung University, Taoyuan, Taiwan5Genomic Medicine Research Core Laboratory, Chang Gung Memorial Hospital, Linkou, Taiwan, Dianjianyi Sun6,Department of Epidemiology and Biostatistics, School of Public Health, Peking University Health Science Center, Beijing, China7Department of Epidemiology, Tulane University School of Public Health and Tropical Medicine, New Orleans, LA 70112, USA, Austin Quach1Department of Human Genetics, David Geffen School of Medicine, University of California Los Angeles, Los Angeles, CA 90095, USA, Alex P. Reiner8Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, WA 98109, USA, Charles Kooperberg8Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, WA 98109, USA, Luigi Ferrucci9Longitudinal Studies Section, Translational Gerontology Branch, National Institute on Aging, National Institutes of Health, Baltimore, MD 21224, USA, Lifang Hou10Center for Population Epigenetics, Robert H. Lurie Comprehensive Cancer Center and Department of Preventive Medicine, Northwestern University Feinberg School of Medicine, Chicago, IL 60611, USA, Andrea A. Baccarelli11Laboratory of Environmental Epigenetics, Departments of Environmental Health Sciences Epidemiology, Columbia University Mailman School of Public Health, New York, NY 10032, USA, Yun Li12Departments of Genetics, Biostatistics, Computer Science, University of North Carolina, Chapel Hill, NC 27599, USA, Sarah E. Harris13,Centre for Cognitive Ageing and Cognitive Epidemiology, Department of Psychology, University of Edinburgh, Edinburgh EH8 9JZ, UK14Department of Psychology, University of Edinburgh, Edinburgh EH8 9JZ, UK, Janie Corley13,Centre for Cognitive Ageing and Cognitive Epidemiology, Department of Psychology, University of Edinburgh, Edinburgh EH8 9JZ, UK14Department of Psychology, University of Edinburgh, Edinburgh EH8 9JZ, UK, Adele Taylor13,Centre for Cognitive Ageing and Cognitive Epidemiology, Department of Psychology, University of Edinburgh, Edinburgh EH8 9JZ, UK14Department of Psychology, University of Edinburgh, Edinburgh EH8 9JZ, UK, Ian J. Deary13,Centre for Cognitive Ageing and Cognitive Epidemiology, Department of Psychology, University of Edinburgh, Edinburgh EH8 9JZ, UK14Department of Psychology, University of Edinburgh, Edinburgh EH8 9JZ, UK, James D. Stewart15Department of Epidemiology, Gillings School of Global Public Health, University of North Carolina, Chapel Hill, NC 27599, USA, Eric A. Whitsel15,Department of Epidemiology, Gillings School of Global Public Health, University of North Carolina, Chapel Hill, NC 27599, USA16Department of Medicine, School of Medicine, University of North Carolina, Chapel Hill, NC 27599, USA, Themistocles L. Assimes17,VA Palo Alto Health Care System, Palo Alto, CA 94304, USA18Department of Medicine, Stanford University School of Medicine, Stanford, CA 94305, USA, Wei Chen7Department of Epidemiology, Tulane University School of Public Health and Tropical Medicine, New Orleans, LA 70112, USA, Shengxu Li19Children’s Minnesota Research Institute, Children’s Hospitals and Clinics of Minnesota, Minneapolis, MN 55404, USA, Massimo Mangino3Department of Twin Research and Genetic Epidemiology, Kings College London, London SE1 7EH, UK, Jordana T. Bell3Department of Twin Research and Genetic Epidemiology, Kings College London, London SE1 7EH, UK, James G. Wilson20Department of Physiology and Biophysics, University of Mississippi Medical Center, Jackson, MS 39216, USA, Abraham Aviv21Center of Development and Aging, New Jersey Medical School, Rutgers State University of New Jersey, Newark, NJ 07103, USA, Riccardo E. Marioni2,Centre for Genomic and Experimental Medicine, Institute of Genetics & Molecular Medicine, University of Edinburgh, Edinburgh EH4 2XU, UK13Centre for Cognitive Ageing and Cognitive Epidemiology, Department of Psychology, University of Edinburgh, Edinburgh EH8 9JZ, UK, Kenneth Raj22Radiation Effects Department, Centre for Radiation, Chemical and Environmental Hazards, Public Health England, Chilton, Didcot, Oxfordshire OX11 0RQ, UK, Steve Horvath1,Department of Human Genetics, David Geffen School of Medicine, University of California Los Angeles, Los Angeles, CA 90095, USA23Department of Biostatistics, Fielding School of Public Health, University of California Los Angeles, Los Angeles, CA 90095, USA
- 1Department of Human Genetics, David Geffen School of Medicine, University of California Los Angeles, Los Angeles, CA 90095, USA
- 2Centre for Genomic and Experimental Medicine, Institute of Genetics & Molecular Medicine, University of Edinburgh, Edinburgh EH4 2XU, UK
- 3Department of Twin Research and Genetic Epidemiology, Kings College London, London SE1 7EH, UK
- 4Department of Biomedical Sciences, Chang Gung University, Taoyuan, Taiwan
- 5Genomic Medicine Research Core Laboratory, Chang Gung Memorial Hospital, Linkou, Taiwan
- 6Department of Epidemiology and Biostatistics, School of Public Health, Peking University Health Science Center, Beijing, China
- 7Department of Epidemiology, Tulane University School of Public Health and Tropical Medicine, New Orleans, LA 70112, USA
- 8Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, WA 98109, USA
- 9Longitudinal Studies Section, Translational Gerontology Branch, National Institute on Aging, National Institutes of Health, Baltimore, MD 21224, USA
- 10Center for Population Epigenetics, Robert H. Lurie Comprehensive Cancer Center and Department of Preventive Medicine, Northwestern University Feinberg School of Medicine, Chicago, IL 60611, USA
- 11Laboratory of Environmental Epigenetics, Departments of Environmental Health Sciences Epidemiology, Columbia University Mailman School of Public Health, New York, NY 10032, USA
- 12Departments of Genetics, Biostatistics, Computer Science, University of North Carolina, Chapel Hill, NC 27599, USA
- 13Centre for Cognitive Ageing and Cognitive Epidemiology, Department of Psychology, University of Edinburgh, Edinburgh EH8 9JZ, UK
- 14Department of Psychology, University of Edinburgh, Edinburgh EH8 9JZ, UK
- 15Department of Epidemiology, Gillings School of Global Public Health, University of North Carolina, Chapel Hill, NC 27599, USA
- 16Department of Medicine, School of Medicine, University of North Carolina, Chapel Hill, NC 27599, USA
- 17VA Palo Alto Health Care System, Palo Alto, CA 94304, USA
- 18Department of Medicine, Stanford University School of Medicine, Stanford, CA 94305, USA
- 19Children’s Minnesota Research Institute, Children’s Hospitals and Clinics of Minnesota, Minneapolis, MN 55404, USA
- 20Department of Physiology and Biophysics, University of Mississippi Medical Center, Jackson, MS 39216, USA
- 21Center of Development and Aging, New Jersey Medical School, Rutgers State University of New Jersey, Newark, NJ 07103, USA
- 22Radiation Effects Department, Centre for Radiation, Chemical and Environmental Hazards, Public Health England, Chilton, Didcot, Oxfordshire OX11 0RQ, UK
- 23Department of Biostatistics, Fielding School of Public Health, University of California Los Angeles, Los Angeles, CA 90095, USA
* Joint last authors
